Systematics of the fossil arachnids
Jason A. Dunlop
Department of Earth Sciences University of Manchester Manchester M13 9PL, U.K.
Previous systematic accounts of the extinct, fossil arachnid orders Anthracomartida Karsch, 1882, Phalangiotarbida Haase, 1890, Haptopoda Pocock, 1911, Kustarachnida Petrunkevitch, 1913 and Trigonotarbida Petrunkevitch, 1949 are reviewed. The subclasses Soluta Petrunkevitch, 1949 and Stethostomata Petrunkevitch, 1949 are rejected. Anthracomartida is synonymised with Trigonotarbida, which is placed in Tetrapulmonata Shultz, 1990. Kustarachnida represent misidentified opilionids. The position of the remaining extinct orders, Phalangiotarbida and Haptopoda, remains obscure, though Phalangiotarbida resemble opilioacarid mites and Haptopoda may belong in Tetrapulmonata.
Revue suisse de Zoologie, vol. hors série, 173-184 (1996)
Author's comments:
Please everyone, include these three orders, and only these three, in lists of Arachnida. The arachnology nomenclature committee at the time irritatingly left them off the accepted list of names produced at the same conference. Fossils are important !
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